Skip to main content Skip to navigation
Hudson Biological Reserve Diversity

Insect and Arthropod Diversity

Dilworth, K. A., Borowiec, M. L., Cohen, A. L., Mickelson, G. S., Oeller, E. C., Crowder, D. W., & Clark, R. E. 2021. Ants of the Palouse Prairie: diversity and species composition in an endangered grassland. Biodiversity data journal, 9, e65768.

Looney, C. 2008. Habitat loss and fragmentation on the Palouse and its impact on arthropod conservation. Ph.D. Dissertation, Department of Environmental Science, University of Idaho.

T.D. Hatten, C. Looney, J. P. Strange, N. A. Bosque-Pérez. 2013. Bumble bee fauna of Palouse Prairie: Survey of native bee pollinators in a fragmented ecosystem, Journal of Insect Science, Volume 13, Issue 1, 26,


Dr. Pam Brunsfeld, retired University of Idaho Systematic Botany Instructor and retired Curator of UI Stillinger Herbarium inspecting a Castilleja sp., likely C. lutescens (Greenm.) Rydb. (Pale paintbrush)

Native Bombus sp. (bumblebee) on Balsamorhiza sagitatta (Arrowleaf Balsamroot)

Selasphorus calliope, a hummingbird native to the Palouse Prairie

Soil Microbial Diversity

(See section below for cyanobacterial and algal diversity)

Daniel C. Schlatter, Christopher M. Baugher, Kendall Kahl, David R. Huggins, Jodi L. Johnson-Maynard, Timothy C. Paulitz. 2019. Bacterial communities of soil and earthworm casts of native Palouse Prairie remnants and no-till wheat cropping systems. Soil Biology and Biochemistry. 139:107625.

Laura L. Ingwell, Christelle Lacroix, Paul R. Rhoades, Alexander V. Karasev, Nilsa A. Bosque-Pérez. 2017. Agroecological and environmental factors influence Barley yellow dwarf viruses in grasslands in the US Pacific Northwest, Virus Research. 241: 185-195.

Westover, K. M., A. C. Kennedy, and S. E. Kelley. 1997. Patterns of rhizosphere microbial community structure associated with co-occurring plant species. Journal of Ecology 85: 863-873.

Mammal Diversity

Randall, J. A. and R. E. Johnson 1979. Population densities and habitat occupancy by Microtus longicaudus and Microtus montanus. Journal of Mammalogy 60: 217-219

Balsamorhiza sagitatta (Arrowleaf Balsamroot) and other native species on Smoot Hill

Arbuscular Mycorrhizal Fungi (AMF) in plant roots (Cheeke Lab, WSU)

Illustration of Thomomys talpoides (Pocket Gopher), native to the Palouse Prairie 

(From Christensen and Larrison, Mammals of the Pacific Northwest: A Pictorial

Moss and Lichen Diversity (Biological Soil Crusts)

(Research in progress)

Cyanobacterial and Algal Diversity (Biological Soil Crusts)

Ashley, J. 1988. Soil algal population and community biology: lLaboratory and field studies. Ph.D. Dissertation, Department of Botany, Washington State University.

Karolina Fučíková, Jared C. Rada & Louise A. Lewis (2011) The tangled taxonomic history of Dictyococcus, Bracteacoccus and Pseudomuriella (Chlorophyceae, Chlorophyta) and their distinction based on a phylogenetic perspective, Phycologia, 50:4, 422-429, DOI: 10.2216/10-69.1

Metting, F. B. 1979. A comparative study of algal communities on cultivated and uncultivated portions of a Schumacher silt loam. Ph.D. Dissertation, Department of Botany, Washington State University.

Metting, F. B. 1980. New species of green microalgae (Chlorophycophyta) from an eastern Washington silt loam. Phycologia 19: 296–306.




Moss, Bryum sp., viewed through a magnifying hand lens

Biological soil crust consisting of mosses (green) and lichens (black and white)

Illustration of algal morphology


(From R. Daubenmire, 1970. Steppe vegetation of Washington. Technical Bulletin 62. Pullman, WA: Washington State University, College of Agriculture, Washington Agricultural Experiment Station.)