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Jon Mallatt




Jon Mallatt photo
Jon Mallatt
Conscious animals of ancient ocean, 330 million years ago, from Feinberg and Mallatt 2016 book
Carboniferous animals said to be conscious are shaded, according to our theory of the evolution of consciousness.
Conscious animals in the Age of reptiles 220 million years ago (shaded) according to theory of Feinberg and Mallatt 2016.
Conscious animals in the Triassic (shaded) according to our theory.

Dr. Mallatt’s CV curriculum vitae






NAME: Jon Moreland Mallatt                                                               DATE: Jan. 29, 2016




Institutions, degrees, dates


9/66-6/70         Whitefish Bay High School; Milwaukee, Wisconsin

B.S.                 9/70-6/74         University of Wisconsin, Zoology

Ph.D.                9/74-12/79       University of Chicago, Department of Anatomy



Positions and dates


Assistant Professor, Departments of Zoology and Basic Medical Sciences, Washington State University, 1/80-8/84

Associate Professor, Departments of Zoology/Biological Sciences and Basic Medical Sciences,

Washington State University, 8/84-present

Affiliate Associate Professor, University of Washington, Department of Biological

Structure, 3/84-present




American Association for the Advancement of Science



WAMI Teaching Award for Excellence in Teaching, Academic Year 1995-96.

WWAMI Teaching Award for Excellence in Teaching, Academic Year 2000-2001

WWAMI Teaching Award for Excellence in Teaching, Academic Year 2003-04

Outstanding Faculty Member at WSU, Interfraternity Council and Panhellenic Association, 2003-04

WSU Mortor Board Distinguished Professor, 2005.

WWAMI Teacher Superior in Perpetuity Award, Academic Year 2011 onward.

Excellent Faculty Senator commendation, Dec. 2013.


INSTRUCTION (1980 to present)


Spring                          Biol 315                 Gross & Microanatomy

Biol 497                 Instructional Practicum


Fall                              Med Sci 510         Histology (50%)

Med Sci 511         Anatomy of the Trunk (50%)




The anatomy textbook that I co-authored (see PUBLICATIONS) is in its seventh edition and is selling well.





  • “Amphioxus, Cambrian fossil animal Haikouella, and the origin of the vertebrates.” At the Symposium on Prevertebrate-Vertebrate Transition (Aspects of Vertebrate Origins). Charles University, Prague, Czech Republic, June 2, 2005. Invited.
  • “How fossils and comparative morphology can inspire Evo-devo studies of the origin of jawed vertebrates.” Symposium on cyclostomes and origin of jawed vertebrates, Kobe, Japan, January 24-25, 2008. Invited.
  • ‘Phylogenetic tree assembled from rRNA sequences from across Metazoa.” Charles D. Walcott and the Discovery of the Burgess Shale: International Conference on the Cambrian Exoplosion. ICCE-Banff, Canada, August 3-8, 2009.


PUBLICATIONS (from 1990 and before)


  1. Mallatt, J. 1979. Journal of Morphology 62:249-273.
  2. Moore, J.W., and J. Mallatt. 1980. Canadian Journal of Fisheries and Aquatic Sciences 37:1658-1664.
  3. Mallatt, J. 1981. Journal of Zoology, London 194:103-142.
  4. Mallatt, J. 1982. Biological Bulletin 163:197-210.
  5. Mallatt, J. 1982. Annals of Science 39:371-380.
  6. Mallatt, J. 1983. Journal of Fish Biology 22:293-301.
  7. Mallatt, J., and R. Ridgway. 1984. Journal of Morphology 180:271-296.
  8. Mallatt, J. 1984. Zoological Journal of the Linnean Society 82:261-272.
  9. Mallatt, J. 1984. Journal of Zoology, London 204:169-183.
  10. Mallatt, J., R. Ridgway, and C. Paulsen. 1986. Canadian Journal of Zoology 63:155-164.
  11. Mallatt, J. 1985. Canadian Journal of Fisheries and Aquatic Sciences 42:630-648.
  12. Mallatt, J. 1985. In: Evolutionary Biology of Primitive Fishes. R.E. Foreman, A.

Gorbman, J.M. Dodd, and R. Olsson, editors. NATO ASI Series A: Life Sciences 103, Plenum, New York. Pp. 59-68.

  1. Mallatt, J., and C. Paulsen. 1985. American Journal of Anatomy 177:243-269.
  2. Mallatt, J., M. G. Barron, and C. McDonough. 1986. Bulletin of Environmental

Contamination and Toxicology 37:281-288.

  1. Mallatt, J., D. M. Conley, and R. L. Ridgway. 1987. Canadian Journal of Zoology 65:1956­1965.
  2. Mallatt, J., and M.G. Barron. 1988. Archives of Environmental Contamination and Toxicology. 17:73-83.
  3. Conley, D.M., and J. Mallatt. 1988. Canadian Journal of Zoology 66:2398-2405.
  4. Stinson, C. M., and J. Mallatt. 1989. Aquatic Toxicology 15: 237-252.
  5. Mallatt, J., and Stinson, C. 1990. Archives of Environmental Contamination and Toxicology. 19:307-313.


PUBLICATIONS (after 1990)


  1. Marieb, E. and J. Mallatt. 1992. Human Anatomy. Benjamin/Cummings Publishing Co.

San Francisco. 800 pages. (Second Edition, 1996, Third edition, 2001, Fourth edition, 2005, Fifth edition, 2008, Sixth Edition, 2011, Seventh Edition, 2014).

  1. Mallatt, J., R. McCall, J. Bailey, and J. Seelye. 1994. Effects of lampricides on the gill

ultrastructure of larval sea lampreys and rainbow trout fry. Canadian Journal of Zoology.


  1. Mallatt, J., J. Bailey, S. Lampa, M. Evans, and W. Tate. 1995. Quantitative ultrastructure

of gill epithelial cells in the larval lamprey Petromyzon marinus. Canadian Journal of

Fisheries and Aquatic Sciences 52:1150-1164.

  1. Mallatt, J., J. Bailey, S. Lampa, M. Evans, and S. Brumbaugh. 1995. A fish-gill system for quantifying the ultrastructural effects of methylmercury, Kepone, and heat shock.

Canadian Journal of Fisheries and Aquatic Sciences 52:1165-1185.

  1. Mallatt, J. 1996. Ventilation and the origin of jawed vertebrates: A new mouth.

Zoological Journal of the Linnean Socity 117:329-404.

6         Mallatt, J. 1997. Shark pharyngeal muscles and early vertebrate evolution. Acta

Zoologica, Stockholm 78:279-294.

  1. Mallatt, J. 1997. Crossing a major morphological boundary: The origin of jaws in

vertebrates. Zoology, Applied and Complex Systems (ZACS) 100:128-140.

  1. Bartels, H., I. C. Potter, K. Niemeier, and J. Mallatt. 1998. Categorisation of mitochondria­ rich cells in the gill epithelium of the freshwater phases in the life cycle of lampreys. Cell and Tissue Research 291 :337-349.
  2. Mallatt, J. and J. Sullivan. 1998. 28S and 18S rDNA sequences support the monophyly of lampreys and hagfishes. Molecular Biology and Evolution 15(12): 1706-1718.
  3. Mallatt, J. 2000. Chordate and vertebrate body structure: Pharyngeal arches and

derivatives. In: Encyclopedia of Paleontology. R. Singer and M. K. Diamond, editors.

Fitzroy Dearborn Publishers. pp.871-877

  1. Hutchinson, M., J. Mallatt, and E. Marieb. 2001. Atlas of the Human Skeleton. Benjamin

Cummings/Addison Wesley Longman

  1. Mallatt, J., J. Sullivan, and C.J. Winchell. 2001. The relationship of lampreys to

hagfishes: A spectral analysis of ribosomal RNA sequences. In: Major Events in Early Vertebrate Evolution: Palaeontology, Phylogeny, and Development. P. Ahlberg, editor. Taylor and Francis, Ltd, pp. 106-118.

  1. Mallatt, J. and C.J. Winchell. 2002. Testing the new animal phylogeny: First use of combined large-subunit and small-subunit rRNA gene sequences to classify the protostomes. Mol. BioI. Evol. 19:289-301.
  2. Winchell, C.J., J. Sullivan, C.B. Cameron, B.J. Swalla, and J. Mallatt. 2002. Evaluating hypotheses of deuterostome phylogeny and chordate evolution with new LSU and SSU ribosomal DNA data. Molecular Biology and Evolution. 19:762-776.
  3. Hutchinson, M., J. Mallatt, and E. Marieb. 2003. A Brief Atlas of the Human Body. Benjamin Cummings/Addison Wesley Longman.
  4. Mallatt, J. and J. Chen. 2003. Fossil sister group of craniates: Predicted and found. Journal of Morphology 258:1-31.
  5. Mallatt,J., J. Chen, and N.D. Holland. 2003. Comment on “A new species of

yunnanozoan with implications for deuterostome evolution.” Science 300:1372c.

  1. Mallatt,J., J.R. Garey, and J.W. Shultz. 2004. Ecdysozoan phylogeny and Bayesian inference; first use of nearly complete 28S and 18S rRNA gene sequences to classify the arthropods and their kin. Molecular Phylogenetics and Evolution. 31:178-191.
  2. Winchell, C.J., A.P. Martin, and J. Mallatt. 2004. Phylogeny of elasmobranchs based on LSU and SSU ribosomal RNA genes. Molecular Phylogenetics and Evolution 31:214-224.
  3. Luan, Y.X., J. Mallatt, R.D. Xie, Y.M. Yang, and W.Y. Yin. 2005. The phylogenetic positions of three basal hexapod groups (Protura, Diplura, and Collembola) based on ribosomal RNA gene sequences. Molecular Biology and Evolution. 22:1579-1592.
  4. Mallatt, J., and G. Giribet. 2006. Further use of nearly complete 28S and 18S rRNA

genes to classify Ecdysozoa: 37 more arthropods and a kinorhynch. Molecular

Phylogenetics and Evolution. 40:772-794.

  1. Mallatt, J., and C.J. Winchell. 2007. Ribosomal RNA genes and deuterostome phylogeny revisited: More cyclostomes, elasmobranchs, reptiles and a brittle star. Molecular Phylogenetics and Evolution. 43:1005-1022.
  2. Mallatt, J. 2008. The origin of the vertebrate jaw: Neoclassical ideas versus newer, development-based ideas. Zoologicial Science 25:990-998.
  3. Mallatt, J. 2009. Evolution and phylogeny of chordates. In: Encylopedia of Neuroscience. M. Binder, H. Hirokawa, and U. Windhorst, editors. Springer-Verlag, pp. 1201-1208.
  4. Mallatt, J., C. Waggoner Craig, and M. Yoder. 2010. Nearly complete rRNA genes assembled from across the metazoan animals: Effects of more taxa, a structure-based alignment, and paired-sites evolutionary models on phylogeny reconstruction. Molecular Phylogenetics and Evolution. 55:1-17.
  5. Mallatt, J., C. Waggoner Craig, and M. Yoder. 2012. Nearly complete rRNA genes from 371 Animalia: Updated structure-based alignment and detailed phylogenetic analysis. Molecular Phylogenetics and Evolution. 64:603-617.
  6. Brunellli, J., J. Mallatt, R. Leary, M. Alfaqih, R. Phillips, G. Thorgaard. 2013. Y chromosome phylogeny for cutthroat trout (Onchorhynchus clarkii) subspecies is generally concordant with those of other markers. Molecular Phylogenetics and Evolution. 66: 592-602.
  7. Mallatt, J., N. Holland. 2013. Pikaia gracilens Walcott: Stem chordate, or already specialized in the Cambrian? Journal of Experimental Zoology 320B: 247-271.
  8. Feinberg, T., J. Mallatt. 2013. The evolutionary and genetic origins of consciousness in the Cambrian Period over 500 million years ago. Frontiers in Consciousness Research Oct. 2013, 4, Article 667.
  9. Mallatt, J., K. Chittenden. 2014. The GC content of LSU rRNA evolves across topological and functional regions of the ribosome in all three domains of life. Molecular Phylogenetics and Evolution 72: 17-30.
  10. Chen, W-J., M. Koch, J. Mallatt, Y-X. Luan. 2014. Comparative analysis of mitochondrial genomes in Diplura (Hexapoda, Arthropoda): Taxon sampling is crucial for phylogenetic inferences. Genome Biology and Evolution 6, 105-120.
  11. Pradel, A., Maisey, J. G., Tafforeau, P., Mapes, R. H., and Mallatt, J. 2014. A Palaeozoic shark with osteichthyan-like branchial arches. Nature, 509 (7502), 608-611.
  12. Feinberg, T. and J. Mallatt. In Press. Neurobiological naturalism. In: Biophysics of Consciousness: A Foundational Approach. R. Poznanski, J. Tuszynski, and T. Feinberg, editors. World Scientific Pub. Co. Publication date: Feb. 21, 2016.
  13. Feinberg, T. and J. Mallatt. In Press. The evolutionary origins of consciousness. . In: Biophysics of Consciousness: A Foundational Approach. R. Poznanski, J. Tuszynski, and T. Feinberg, editors. World Scientific Pub. Co. Publication date: Feb. 21, 2016.
  14. Feinberg, T. and J. Mallatt. In Press. The Ancient Evolutionary Origins of Consciousness: How the Brain Created Experience. MIT Press. Publication date: March 25, 2016.