Publications
- Robb, B.C., S. J. Galla, T. T. Caughlin, J. J. Mitchell, P. J. Olsoy, J. D. Nobler, M. R. Fremgen-Trantino, D. M. Delparte, L. A. Shipley, J. L. Rachlow, C. Dadabay, and J. S. Forbey. In review. Near infrared spectroscopy predicts concentrations of phytochemicals and browsing by vertebrate herbivores. PNAS (March 2025).
- Burgstahler*, K., L. A. Shipley, A. Stift, S. Woodley, and D. Thornton. In review. Effectsof wildfire on the distribution of mammals and birds in a working shrub-steppe landscape. Fire Ecology (March 2025).
- Huggler*, K. S., R. A. Smiley*, B. L. Wagler, A. B. Courtemanch, Z. Gregory, K. L. Monteith, L. A. Shipley, C. Steward, P. Wik, E. F. Cassirer, and R. A. Long. In review. Should I stay or should I go. Causes and consequences of intraspecific variation in site fidelity. Movement Ecology (February 2025).
- Torres Ferreira*, A., L. A. Shipley, A. Stahl, M. C. Camp, and A. Meddens. In review. Handheld photogrammetry advances capabilities to systematically characterize broken-rock habitat. Wildlif Society Bulletin (March 2025).
- Anderson*, K.A., L. A. Shipley, S. Galla, M. Calahan, S. Berry, S. Hudon, and J. S. Forbey. In review. Food or family? How gut microbes respond to diet and phylogeny in tow deer species. Ecosphere (February 2025).
- Balluffi-Fry*, J., L. A. Shipley, R. Zijlstra, E. Bork, M. Humphries, and S.Boutin. In review. Nutrient balancing by a wild browsing herbivore: nutritional geometry of snowshoe hares (Lepus americanus). Journal of Animal Ecology (February 2025).
- Hellesto*, R., L. A. Shipley, P. Bennett**, K. Huggler*, and M. Devivo. In revision. Early fawn-rearing habitat of mule deer in an agricultural landscape. Ecosphere (March 2024)
- Huggler*, K. S., R. A. Smiley*, B. L. Wagler, A. B. Courtemanch, Z. Gregory, D. Lutz, K. L. Monteith,
- A. Shipley, C. Stewart, P. Wik. E. F. Cassirer, and R. A. Long. In review. Causes and
consequences of life-history variation in a long-lived mammal. Wildlife Monographs (October
2024).
- Monzingo*, D. S., J. G. Cook, R. C. Cook, L. Irwin, and L. A. Shipley. In revision. Dynamics of
Forage Resources for Elk in North-Central Idaho: 1870-2015. Journal of Wildlife
Management (January 2025).
- Celebrezze, J. V., M. A. Okikiola*, D. A. Glavich, L. A. Shipley, and A. J. H. Meddens. 2025.
Classifying rocky land cover using random forest modeling: lessons learned and potential
applications in Washington, USA. Remote Sensing 17/915.
- Hellesto*, R., L. A. Shipley, and R. A. Long. 2025. Seasonal habitat selection and migration of
mule deer in an agricultural landscape. Journal of Wildlife Management . Journal of Wildlife
Management https://doilrog/10.1002/jwmg.70014..
- Robatchek*, S., C. White, R. Long, and L. A. Shipley. 2025. Using dynamic foodscape models
to assess bottom-up constraints on population performance of herbivores. Ecological
Applications doi:10.1002/eap.70015.
- Bilodeau-Hussey*, N. M., K. S. Huggler*, E. F. Cassirer, H. Miyasaki, M. Hurley, S. Roberts, L. A.
Shipley, and R. A. Long. 2025. Effects of maternal condition, disease status, and behavior
on survival of juvenile bighorn sheep. Journal of Wildlife Management
https://doi.org/10.1002/jwmg.22721
- Lopez*, K., K. Huggler*, D. Jackson, L. A., Shipley and R. A. Long. 2024. Linking summer
nutrition to behavior and performance of black-tailed deer. The Journal of Wildlife Management
2024:e22679.
- Whiles*, L., L. A. Shipley, J. R. Akins, R. G. Christophersen, J. I. Ransom, T. Levi, and D. H.
Thornton. 2024. Predation risk for hoary marmots in in Washington’s North Cascades Mountains.
Journal of Mammalogy gyae086, https://doi.org/10.1093/jmammal/gyae086
- Ganz*, T. R., S. B. Bassing, M. DeVivo, B. Gardner, B. Kertson, L. Satterfield, L. A. Shipley, B.
Turnock, S. L. Walker, D. Abrahamson, A. Wirsing, and L. Prugh. 2024. White-tailed deer
population dynamics in a multi-predator landscape shaped by humans. Ecological Applications
DOI: 10.1002/eap.3003
- Peziol*, M., M. L. Elbroch, L. A. Shipley, D. R. Evans, H. Quigley, and D. Thornton. 2023.
Large carnivore foraging contributes to heterogeneity in nutrient cycling. Landscape Ecology
38:1497-1509.
- Monzingo*, D. S., J. G. Cook, R. C. Cook, J. S. Horne and L. A. Shipley. 2023. Influences of
succession and biogeoclimate on forage Resources for elk in Northern Idaho. Northwest
Science 96:94-1116.
- Cook, R. C., L. A. Shipley, J. G. Cook, M. J. Camp, D. S. Monzingo*, S. L. Robatcek, S. L. Berry, I. T.
Hull*, W. L. Myers, K. A. Denryter, and R. A. Long. 2022. Sequential detergent fiber assay
results used for nutritional ecology research: Evidence of bias since 2012. Wildlife Society
Bulletin 46:e1348.
- Konôpka, B., V. Šebeň, J. Pajtík, and L. A. Shipley. 2022. Influence of tree species and size on
bark browsing by large wild herbivores. Plants 11:2925, https://doi.org/10.3390/plants11212925.
- Olsoy*, P. J., C. R. Milling*, J. D. Nobler*, M. J. Camp*, L. A. Shipley, J. S. Forbey, J. L. Rachlow,
and D.H. Thornton. 2022. Food quality, security, and thermal refuge influence the use of microsites and patches by pygmy rabbits (Brachylagus idahoensis) across landscapes and
seasons Ecology and Evolution 12:e8892 DOI: 10.1002/ece3.8892.
- Monzingo*, D. S., L. A. Shipley, R. C. Cook, and J. G. Cook. 2022. Factors influencing
predictions of understory vegetation biomass from visual cover estimates. Wildlife Society
Bulletin 46:e1300, https://doi.org/10.1002/wsb.1300.
- Gowda, J. H., M. Blackhall, L. A. Shipley, T. Kitzberger, and F. Tiribelli. 2022. Are digestibility
and flammability related? Two variables shaping landscape dynamics of Northwestern
Patagonian forests. Forest Ecology and Management 503:119810.https://doi.org/10.1016/j.foreco.2021.119810
- Robb*, B., P. J. Olsoy*, J. J. Mitchell, T. T. Caughlin*, D. M. Delparte, M. R. Fremgen-Tarantino, J. D.
Nobler*, J. L. Rachlow, L. A. Shipley, and J. S. Forbey. 2021. Near-infrared spectroscopy
aids ecological restoration by classifying variation of taxonomy and phenology of a native shrub.
Restoration Ecology 30(5) e13584:, doi: 10.1111/rec.13584.
- Staudenmaier*, A. R., L. A. Shipley, M. J. Camp, J. S. Forbey, A. E. Hagerman, A. E. Brandt**, and
- H. Thornton. 2021. Mule deer do more with less: comparing their nutritional requirements
and tolerances with white-tailed deer. Journal of Mammalogy 103:178-195,
https://doi.org/10.1093/jmammal/gyab116
- Staudenmaier*, A. R., L. A. Shipley, A. Bibelnieks*, M.J. Camp, and D. H. Thornton. 2021.
Habitat use and spatio-temporal interactions of mule and white-tailed deer in an area of sympatry
in NE Washington. Ecosphere 12(11):e03813.
- Konôpka, B., V. Šebeň, J. Pajtík, and L. A. Shipley. 2021. Excluding large wild herbivores
reduced Norway spruce dominance and supported tree species richness in a young, naturally
regenerated stand. Forests 12, 737, https://doi.org/10.3390/f12060737.
- Ulappa*, A. C., L. A. Shipley, R. C. Cook, J. G. Cook, and M. E. Swanson. 2020. Silvicultural
herbicides and forest succession influence understory vegetation and nutritional ecology of
black-tailed deer in managed forests. Forest Ecology and Management,
https://doi.org/10.1016/j.foreco.2020.118216
- Merems*, J. L., L. A. Shipley, T. Levi, J. Ruprecht, D. A. Clark, M. J. Wisdom, N. J. Jackson, K. M
Stewart, and R. A. Long. 2020. Nutritional-landscape models link habitat use to condition of
Mule deer (Odocoileus hemionus). Frontiers in Ecology and Evolution 8:98. doi:
10.3389/fevo.2020.00098
- Camp, M. J., L. A. Shipley, J. Varner, and B. D. Waterhouse. 2020. Activity patterns and foraging
behavior of American pikas (Ochotona princeps) differ between Craters of the Moon and alpine
talus in Idaho. Western North American Naturalist 80:49-69.
- Olsoy*, P.J., J. S. Forbey, L. A. Shipley, J. L. Rachlow, B. C. Robb**, J. D. Nobler*, and D. H.
Thornton. 2020. Mapping foodscapes and sagebrush morphotypes with unmanned aerial
Systems for multiple herbivores. Landscape Ecology 35:921-936.
- Hull, I. T.*, L. A. Shipley, S. L. Berry*, C. Loggers, and T. R. Johnson. 2020. Effects of fuel
reduction timber harvests on forage resources for deer in northeastern Washington. Forest
Ecology and Management 458:117757
- Konôpka, B., J. Pajtík, M. Bošeľa, Vladimír Šebeň, and L. A. Shipley. 2020. Modeling forage
potential for red deer (Cervus elaphus) – a tree level approach. European Journal of Forest
Research 139:419-430.
- Schrempp, T.*, J. L. Rachlow, T. R. Johnson, L. A. Shipley, R. Long, J. Aycrigg, and M. Hurley.
- Linking forest management to moose population trends: The role of the nutritional
landscape. PLOS ONE https://doi.org/10.1371/journal.pone.0219128
- Berry, S. L.*, L. A. Shipley, R. A. Long and C. Loggers. 2019. Differences in dietary niche and
foraging behavior by sympatric mule and white-tailed deer in Washington. Ecosphere 10:e02815.
- Ebenhoch, K., D*. Thornton, L. A. Shipley, J. Manning, and K. White. 2019. Comparing population
vital rates of resident and translocated greater sage-grouse in Yakima, WA. Journal of Wildlife
Management 83:1314-1325.
- Hull, I. T*., and L. A. Shipley. 2019. Testing the ability of airborne LiDAR to measure forage
resources for wild ungulates in conifer forests. Journal of Forestry 117:492-503 https://doi.org/10.1093/jofore/fvz040 .
- Nobler, J. D.*, M. J. Camp*, M. M. Crowell*, L. A. Shipley, C. Dadabay, J.L. Rachlow, L. James**,
and J. S. Forbey. 2018. Preferences of specialist and generalist mammalian herbivores for
mixtures versus individual plant secondary metabolites. Journal of Chemical Ecology
https://doi.org/10.1007/s10886-018-1030-5
- Konôpka, B., J. Pajtík, and L. A. Shipley. 2018. Intensity of red deer browsing on young rowans
differs between freshly-felled and standing individuals. Forest Ecology and Management
429:511-519.
- Milling, C.R.*, J.L. Rachlow, P.J. Olsoy*, M.A. Chappell, T.R. Johnson, J.S. Forbey, L.A. Shipley, and D.H.Thornton. 2018. Habitat structure modifies microclimate: an approach for mapping fine-
scale thermal heterogeneity. Methods in Ecology and Evolution 9:1648-1657.
- Crowell, M. M*., L. A. Shipley, J. S. Forbey, J. L. Rachlow, and R. G. Kelsey. 2018. Dietary partitioning of toxic leaves and fibrous stems differs between sympatric specialist and generalist mammalian herbivores. Journal of Mammalogy 99:565-577, https://doi.org/10.1093/jmammal/gyy018.
- Milling*, C. R., J. L. Rachlow, M. A. Chappell, M. J. Camp*, T. R. Johnson, D. R. Paul, L. A. Shipley,
and J. S. Forbey. 2018. Seasonal temperature acclimatization in a semi-fossorial mammal and
the role of burrows as thermal refugia. PeerJ 6:e4511; DOI 10.7717/peerj.4511.
- Camp*, M. J., L. A. Shipley, C. R. Milling*, J. L. Rachlow, and J. S. Forbey. 2018. Interacting effects of ambient temperature and food quality on the foraging ecology of small mammalian herbivores. Journal of Thermal Biology 71:83-90.
- McMahon, L. A.*, J. L. Rachlow, L. A. Shipley, J. S. Forbey, and T. R. Johnson. 2017. Habitat selection differs across hierarchical behaviors: Selection of patches and intensity of patch use. Ecosphere 8(11):0.1993.
- Olsoy, P, J.*, L. A. Shipley, J. L. Rachlow, J. S. Forbey, N. Glenn, M. Burgess, and D. Thornton.
- Unmanned aerial systems measure structural habitat features for wildlife across
multiple scales. Methods in Ecology and Evolution 9:594-604, DOI:10.1111/2041-210X.12919.
- Camp*, M. J., L. A. Shipley, T. R. Johnson, P. Olsoy*, J. S. Forbey, J. L. Rachlow, and D. Thornton. 2017. The balancing act of foraging: Mammalian herbivores trade-off multiple risks when selecting food patches. Oecologia 185:537-549, DOI 10.1007/500442-017-3957-6.
- Milling*, C. R., J. L. Rachlow, T. R. Johnson, J. S. Forbey, and L. A. Shipley. 2017. Seasonal
variation in behavioral thermoregulation and predator avoidance in a small mammal. Behavioral Ecology 28:1236-1247.
- Forbey, J. S., G. L. Patricelli, D. M. Delparte, A. H. Krakauer, P. J. Olsoy*, M. R. Fremgen*, J. D. Nobler*, L. P. Spaete, L. A. Shipley, J. L. Rachlow, A. K. Dirksen, A. Perry, B. A. Richardson, and N. F. Glenn. 2017. Emerging technology to measure habitat quality and behavior of grouse: examples from studies of greater sage-grouse. Wildlife Biology DOI: 10.2981/wlb.00238
- McMahon*, L. A, J. L. Rachlow, L. A. Shipley, J. S. Forbey, T. R. Johnson, and P. J. Olsoy*. 2017. Evaluation of micro-GPS receivers for tracking small-bodied mammals. PloS ONE 12(3): e0173185. doi:10.1371/journal.pone.0173185
- King*, T. W., R. Salom-Pérez, L. A. Shipley, H. B. Quigley, and D. H. Thornton. 2017. Ocelot latrines: communication centers for Neotropical mammals. Journal of Mammalogy 98:106-113.
- Olsoy*, P. J., T. G. Griggs, A. C. Ulappa*, K. Gehlken*, L. A. Shipley, G. E. Shewmaker, and J. S.
Forbey. 2016. Nutritional analysis of sagebrush by near-infrared spectroscopy. Journal of Arid
Environments 134:125-1131.
- Utz*, J. L., L. A. Shipley, J. L. Rachlow, T. L. Johnstone-Yellin, M. J. Camp*, and J.S. Forbey. 2016.
Understanding tradeoffs between predation and food risks in a specialist mammalian herbivore.
Wildlife Biology 22:167-173.
- Crowell*, M. M., L. A. Shipley, M. J. Camp*, J. L. Rachlow, J. S. Forbey, and T. R.Johnson. 2016.
Selection of food patches by sympatric herbivores in response to concealment and distance from
a refuge. Ecology and Evolution DOI: 10.1002/ece3.1940.
- Camp*, M. J., L. A. Shipley, T. R. Johnson, M. M. Crowell*, J. S. Forbey, and J. L. Rachlow. 2015.
Modeling tradeoffs between plant fiber and toxins: A framework for quantifying risks perceived by
foraging herbivores. Ecology. 96:3292-3302
- Stonehouse*, K. F., L. A. Shipley, J. Lowe, M. T. Atamian, M. E. Swanson, and M. A. Schroeder.
- Habitat selection by sympatric greater sage-grouse and Columbian sharp-tailed grouse.
Journal of Wildlife Management 79:1308-1326.
- Olsoy*, P. J., J. S. Forbey, J. L. Rachlow, J. D. Nobler*, N. F. Glenn, and L. A. Shipley. 2015.
Fearscapes: mapping functional properties of cover for prey with terrestrial LiDAR. Bioscience
65:74-80.
- Rachlow, J. L., R. M. Peter**, L. A. Shipley, and T. R. Johnson. 2014. Sublethal effects of capture
and collaring on wildlife: experimental and field evidence. Wildlife Society Bulletin 38:458-465.
- Ulappa*, A. C., R. G. Kelsey, G. G. Frye*, J. L. Rachlow, L. A. Shipley, X. Pu, L. Bond, and J.
Forbey. 2014. Plant protein and secondary metabolites influence diet selection in a mammalian
specialist herbivore. Journal of Mammalogy 95:834-842.
- Camp*, M. J., J. L. Rachlow, L. A. Shipley, T. R. Johnson, and K. D. Bockting. 2014. Grazing in
Sagebrush rangelands in western North American: Implications for habitat quality for a sagebrush
specialist, the pygmy rabbit. The Rangeland Journal 36:151-159.
- Elias*, B. A., L. A. Shipley, S. McCusker*, R. D. Sayler, and T. R. Johnson. 2013. Effects of genetic
management on reproduction, growth, and survival in captive endangered pygmy rabbits
(Brachylagus idahoensis). Journal of Mammalogy 94:1282-1292.
- Ellsworth*, E., A. J. Wirsing, L. A. Shipley, and D. L. Murray. 2013. Do measures of plant intake and
digestibility from captive feeding trials align with foraging patterns of free-ranging snowshoe
hares? Wildlife Research 40:340-357.
- Scarlata*, C. D., B. A. Elias*, J. R. Godwin, R. A. Powell, D. Shepherdson, L. A. Shipley, and J. L.
Brown. 2013. Influence of environmental conditions and facility on faecal glucocorticoid
concentrations and in captive pygmy rabbits. Animal Welfare 22:357-368.
- Cook*, R. C., J. G. Cook, D. J. Vales, B, K. Johnson, S. M. McCorquodale, L. A. Shipley, R. A. Riggs,
L.L. Irwin, S. L. Murphie, B. L. Murphie, K. A. Schoenecker, F. Geyer, P. B. Hall, R. D. Spencer,
D.A. Immell, D. H. Jackson, B. L. Tiller, P. J. Miller, and L. Schmitz. 2013. Regional and
seasonal patterns of nutritional condition and reproduction in elk. Wildlife Monographs 184.
- Wagoner*, S. J., L. A. Shipley, R. C. Cook*, and L. Hardesty. 2013 Spring cattle grazing and mule
deer nutrition in a bluebunch wheatgrass community. Journal of Wildlife Management 77:897-907.
- Camp*, M. J., B. A. Woods*, J. L. Rachlow, T. R. Johnson, and L. A. Shipley. 2013. Examining the
functional components of cover: The relationship between concealment and visibility in shrub-
steppe habitat. Ecosphere 4:1-14.
- Camp*, M. J., J. L. Rachlow, B. A. Woods*, T. R. Johnson, and L. A. Shipley. 2012. When to run
and when to hide: The influence of concealment, visibility, and proximity to refugia on perception
of risk. Ethology 118:1010-1017.
- Shipley, L. A., E. M. Davis, L. A. Felicetti, S. McClean, and J. S. Forbey. 2012. Mechanisms for
eliminating monoterpenes of sagebrush by specialist and generalist rabbits. Journal of Chemical Ecology 38:1178-1189.
- Scarlata*, C. D., B. A. Elias, J. R. Godwin, R. A. Powell, D. Shepherdson, L. A. Shipley and J. L.
Brown. 2012. Relationship between fecal hormone concentrations and reproductive success in
captive pygmy rabbits (Brachylagus idahoensis). Journal of Mammalogy 93:759-770.
- Scarlata*, C. D., B. A. Elias, J. R. Godwin, R. A. Powell, D. Shepherdson, L. A. Shipley and J. L.
Brown. 2011. Characterizing gonadal and adrenal activity by fecal steroid analysis in pygmy
rabbits (Brachylagus idahoensis). General and Comparative Endocrinology 171:373-380.
- Tollefson*, T. L., L. A. Shipley, W. L. Myers, and N. Dasgupta. 2011. Forage quality’s influence on
Mule deer fawns. Journal of Wildlife Management 75:919-928.
- McCusker*, S., L. A. Shipley, T. N. Tollefson*, M. Griffin, and E. A. Koutsos. 2011. Effects of starch
and fibre in pelleted diets on nutritional status of mule deer (Odocoileus hemionus) fawns. Journa
of Animal Physiology and Animal Nutrition 95: 489 – 498.
- Dungan*,J. D., L. A. Shipley, and R. G. Wright. 2010. Activity patterns, foraging ecology
and summer range carrying capacity of moose in Rocky Mountain National Park, Colorado,
Alces 46: 71-87.
- Tollefson*, T. L., L. A. Shipley, W. L. Myers, D. H. Keisler, and N. Dasgupta. 2010.
The influence of summer and autumn nutrition on body condition and reproduction in lactating
mule deer. Journal of Wildlife Management 74:974-986.
- Lopez-Perez*, E., L. A. Shipley, and W. Myers. 2010. Blood selenium levels of mule deer in eastern
Washington. Journal of Dairy Science 93:125-126.
- Shipley, L. A. 2010. Fifty years of food and foraging in moose: Lessons in ecology from a model
herbivore. Alces 46:1-13.
- McClure, M. F., and L. A. Shipley. 2009. Animal response to nested self-similar patches: A test with
woolly bears. Oikos 118:653-662.
- Johnstone-Yellin*, T. L., L. A. Shipley, W. L. Myers, and H. S. Robinson*. 2009. To twin or not to
twin? Tradeoffs in litter size and fawn survival in mule deer. Journal of Mammalogy 90:453-460.
- Shipley, L. A., J. S. Forbey, and B. D. Moore. 2009. Revisiting the dietary niche: when
is a mammalian herbivore a specialist? Integrative and Comparative Biology 49:274-290.
- Kendrick*, E., L. A. Shipley, A. E. Hagerman, and L. M. Kelly*. 2009. Fruit and fibre: The nutritional
value of figs for a small tropical ruminant, the blue duiker (Cephalophus monticola). Journal of
African Ecology 47:556-566.
- DeGabriel*, J. L., B. D. Moore, L. A. Shipley, A. K. Krockenberger, I. R. Wallis, C. N. Johnson, and W.
- Foley. 2009. Inter-population differences in the tolerance of a marsupial folivore to plant
secondary metabolites. Oecologia 161:534-548.
- Shipley, L. A. 2008. Pygmy rabbits in peril in the U.S.A.
http://www.actionbioscience.org/biodiversity/shipley.html
- Thines*, N. J., L. A. Shipley, J. H. Bassman, J. R. Slusser, and W. Gao. 2008. UV-B effects on
nutritional chemistry of plants and responses of a mammalian herbivore. Oecologia 156:125-135.
- Shipley, L. A. 2007. The influence of bite size on foraging at larger spatial and temporal scales by
mammalian herbivores. Oikos 116:1964-1974.
- Robbins, C. T., J. K. Fortin*, K. D. Rode, S. D. Farley, and L. A. Shipley. 2007. Optimizing protein
intake as a foraging strategy to maximize mass gain in an omnivore. Oikos 116:1675-1682.
- Cook, R. C., T. R. Stephenson, W. L. Myers, J. G. Cook, and L. A. Shipley. 2007. Validating
predictive models of nutritional condition for mule deer. Journal of Wildlife Management
71:1934-1943.
- Thines*, N. J., L. A. Shipley, J. H. Bassman, J. K. Fellman, D. S. Mattison, and J. R. Slusser. 2007.
Effects of enhanced UV-B radiation on plant chemistry: Nutritional consequences for a specialist
and generalist lagomorph.Journal of Chemical Ecology 33:1025-1039.
- Shipley, L. A., T. B. Davila*, N. J. Thines*, and B. A. Elias*. 2006. Nutritional requirements and diet
choices of the pygmy rabbit (Brachylagus idahoensis): A sagebrush specialist. Journal of
Chemical Ecology 32:2455-2474.
- Elias*, B. A., L. A. Shipley, R. D. Sayler and R. S. Lamson*. 2006. Mating behavior and parental
Care In captive pygmy rabbits. Journal of Mammalogy 87:921-928
- Johnstone-Yellin*, T. L., L. A. Shipley, and W. L. Myers. 2006. Evaluating the performance of vaginal
implant transmitters for capturing neonatal mule deer fawns. Wildlife Society Bulletin 34:338-
344.
- Searle*, K., T. Vandervelde*, N. T. Hobbs, L. A. Shipley, and B. A. Wunder. 2006. Spatial Context
influences patch residence time in foraging hierarchies. Oecologia 148:710-719.
- Searle*, K., N. T. Hobbs, B. A. Wunder, and L. A. Shipley. 2006. Preferences in a patchy
landscapes: the influence of scale-specific intake rates and variance in reward. Behavioral
Ecology 17:315-323.
- Searle*, K. R., N. T. Hobbs, and L. A. Shipley. 2005. Should I stay or should I go? Patch departure
decisions by herbivores at multiple scales.Oikos 111:417-424.
- Searle*, K. R., T. Vandervelde*, N. T. Hobbs, and L. A. Shipley. 2005. Gain functions for large
herbivores: tests of alternative models. Journal of Animal Ecology 74:181-189.
- Thines* N. J., L. A. Shipley, and R. D. Sayler. 2004. Effects of cattle grazing on ecology and
habitat of Columbia Basin pygmy rabbits (Brachylagus idahoensis). Biological
Conservation 119:525-534.
- Sponheimer, M., T. Robinson, L. Ayliffe, B. Passey, B. Roeder, L. Shipley, E. Lopez*, T. Cerling, D.
Dearing* and J. Ehleringer. 2003. An experimental study of carbon-isotope fractionation
between diet, hair, and feces of mammalian herbivores. Canadian Journal of Zoology 81:871-
- Felicetti*, L.A., C.T. Robbins, and L.A. Shipley. 2003. Dietary protein content alters energy
expenditure and composition of the gain in grizzly bears (Ursus arctos horribilis).
Physiological and Biochemical Zoology 76:256-261.
- Hobbs, N. T., J. E. Gross, L. A. Shipley, D. E. Spalinger, and B. A. Wunder. 2003. Herbivore
functional response in heterogeneous environments: A contest among models. Ecology 84:666-
681
- Shipley, L. A., and L. A. Felicetti*. 2002. Nitrogen and fiber digestibility of natural forages by blue
duikers. Zoo Biology 21:123-134.
- Nolte, D., L. A. Shipley, and J. Schafer. 2001. Evaluating Wolfin as a repellent to deer on roads in
Washington. Western Journal of Applied Forestry 16:182-186.
- Fletcher*, J. D., L. A. Shipley, W. J. McShea, and D. L. Shumway. 2001. Wildlife herbivory and rare
plants: The effects of white-tailed deer, rodents, and insects on growth and survival of Turk’s
cap lily. Biological Conservation 101:229-238.
- Rode*, K. D., C. T. Robbins, L. A. Shipley, and T. A. Hanley. 2001. The constraints on herbivory by
grizzly bears. Oecologia 128:62-71.
- Fletcher*, J. D., W. J. McShea, L. A. Shipley, and D. Shumway. 2001. The utility of using common
forest forbs to measure browsing pressure by white-tailed deer (Odocoileus virginianus).
Natural Areas Journal 54:413-419.
- Laca, E., L. A. Shipley, and E. Reid*. 2001. Structural anti-quality characteristics of range and
pasture plants. Journal of Range Management 54:413-419.
- Felicetti*, L. A., L. A. Shipley, G. Witmer, and C. T. Robbins. 2000. Digestive strategies of North
American porcupines: Nitrogen, dry matter, and energy digestibility of natural forages.
Biochemical and Physiological Zoology 73:772-780.
- Burton*, N. M., L. A. Shipley, K. Byrne, J. Vierck, and M. V. Dodson. 2000. Isolation and culture
of wapiti (Cervus elaphus) satellite cells. Canadian Journal of Animal Science
80:303-309.
- Brown, W. K., W. K. Hall, L. R. Linton, R. E. Huenefeld*, and L. A. Shipley. 2000.The repellency of 3
compounds to caribou. Wildlife Society Bulletin 28:365-371.
- Wenninger*, P. S., and L. A. Shipley. 2000. A comparison of harvesting, rumination, digestion,
rumination, and passage on fruit, forb, and leaf diets by blue duikers. Oecologia. 123:466-474.
- Shipley, L. A., A. W. Illius, K. Danell, N. T. Hobbs, and D. E. Spalinger. 1999. Predicting bite size
selection of mammalian herbivores: a test of a general model of diet optimization. Oikos 84:55-
- Shipley, L. A., S. Blomquist*, and K. Danell. 1998. Diet choices by free-ranging moose in relation to
plant distribution, chemistry, and morphology in northern Sweden. Canadian Journal of Zoology
76:1722-1733.
- Spalinger, D. E., S. M. Cooper, D. Martin, and L. A. Shipley*. 1997. Is social learning an important
influence on foraging behavior in white-tailed deer? Journal of Wildlife Management 61:611-
621.
- Shipley*, L. A., D. E. Spalinger, J. E. Gross, N. T. Hobbs, and B. A. Wunder. 1996. The dynamics
and scaling of foraging velocity and encounter rate in mammalian herbivores. Functional Ecology
10:234-244.
- Shipley*, L. A., and D. E. Spalinger. 1995. Influence of size and density of browse patches on intake
rates and foraging decisions of young moose and white-tailed deer. Oecologia 104:112-121.
- Shipley*, L. A., J. E. Gross, D. E. Spalinger, N. T. Hobbs, and B. A. Wunder. 1994. The scaling of
intake rate of mammalian herbivores. American Naturalist 143:1055-1082.
- Gross, J. E., L. A. Shipley*, N. T. Hobbs, D. E. Spalinger, and B. A. Wunder. 1993. Functional
response of herbivores in food-concentrated patches: tests of a mechanistic model. Ecology 74:
778-791.
- Shipley*, L. A., and D. E. Spalinger. 1992. Mechanics of browsing in dense food patches: effects
of plant and animal morphology on intake rate. Canadian Journal of Zoology 70:1743-1752.