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Dr. Dan Thornton's Mammal Spatial Ecology and Conservation Lab


PUBLICATIONS (PEER-REVIEWED) **=senior authorship on publication, 1=supervised graduate student, 2= supervised undergraduate student


Thornton, D.H., and D.L. Murray. (submitted). Modeling the historic range of at-risk species to help inform conservation planning: Canada lynx in the contiguous United States. Biological Conservation.


 Gump1, K., and Thornton, D.H.**. 2023.  Trucks vs. Treks: The relative impact of motorized vs. non-motorized recreation on a mammal community. Ecological Applications. e2916.

Anderson1, A., Waller, J., and Thornton, D.H.**. 2023. Partial COVID-19 closure of a national park reveals negative influence of low-impact recreation on wildlife spatiotemporal ecology. Scientific Reports. 13:687.

Anderson1, A., Waller, J., and Thornton, D.H.**. 2023. Canada lynx occupancy and density in Glacier National Park. Journal of Wildlife Management 87:e22383

Peziol1, M., Elbroch, L.M., Shipley. L.S., Evans, R.D., Quigley, H., Thornton, D.H**. 2023. Mountain lion foraging contributes to nutrient cycling. Landscape Ecology 38: 1497-1508.

Lyons, A. Gaines, W., Lewis, J., Maletzke, B., Werntz, D., Thornton, D., Begley, J., Vanbianchi, C., King, T., Blatz, G., and Fitkin, S. 2023. Climate change, wildfire, and forest management challenge survival of Canada lynx in the North Cascades, USA. Journal of Wildlife Management 87:e22410



Olsoy1, P, Milling, C.R., Nobler, J.D., Camp, M.J., Shipley, L.A., Forbey, J.S., Rachlow, J.L., and D.H. Thornton**. 2022. Food quality, security, and thermal refuge influence the use of microsites and patches by pygmy rabbits (Brachylagus idahoensis) across landscapes and seasons. Ecology and Evolution 12:e8892.

Staudenmaier1, A., Shipley, L., Camp, M., Forbey, J., Hageman, A., Brandt, A., and D.H. Thornton. 2022.  Mule deer do more with less: comparing their nutritional requirements and tolerances with white-tailed deer. Journal of Mammalogy 103:178-195.

Jensen1, P, Wirsing, A., and D.H. Thornton**.2022. Using camera-traps to estimate density of snowshoe hare (Lepus americanus): a keystone boreal forest herbivore. Journal of Mammalogy 103:693-710.

 Arrais, R., Widmer, C., Murray, D., Thornton, D.H., and F. Azevedo. 2022. Estimating density of ocelots (Leopardus pardalis) in the Atlantic Forest using spatial and close capture-recapture models. Journal of Mammalogy 103: 1327-1337.

Calderón, A.P., Louvier, J., Planillo, A., Araya-Gamboa, D., Arroyo-Arce, S., Barrrantes-Nunez, M., Carazo-Salazar, J., Corrales-Gutierrez, D., Doncaster, C.P., Foster, R., Garcia, M.J., Garcia-Anleu, R., Harmsen, B., Hernandez-Potosme, S., Leonardo, R., Trigueros, D., McNab, R., Meyer, N., Moreno, R., Salom-Perez, R., Sauma Rossi, A., Thomson, I., Thornton, D., Yahaira, U., Grimm, V., Kramer-Schadt, S. 2022. Occupancy models reveal potential of conservation prioritization for Central American jaguars. Animal Conservation 25:680-691.

Kays, R., Lasky, M., Allen, M.L., Dowler, R.C., Hawkins, M.T.R., Hope, A.G., Kohli, B.A., Mathis, V.L., McLean, B., Olson, L.E., Thompson, C.W., Thornton, D., Widness, J., Cove, M.V. 2022. Which mammals can be identified from camera traps and crowd-sourced photographs? Journal of Mammalogy 103: 767-775. (Editor’s choice)



Staudenmaier1, A., Shipley, L., Bibelmieks, A., Camp, M., and D.H. Thornton**. 2021. Habitat use and spatio-temporal interactions of mule and white-tailed deer in an area of sympatry in NE Washington. Ecosphere 12:e03813

Jensen1, P.O, Meddens, A.J.H., Fisher, S., Wirsing, A.J., Murray, D.L., and D.H. Thornton**. 2021. Broaden your horizon: The use of remotely sensed data for modeling populations of forest species at landscape scales. Forest Ecology and Management 500:119640.

King1, T., Vynne, C., Miller, D., Fisher, S., Fitkin, S., Rohrer, J., Ransom, J., and D.H. Thornton**. 2021. The influence of spatial and temporal scale on the relative importance of biotic vs. abiotic factors for species distributions. Diversity and Distributions 27:327-343

Perera1, L., Garcia-Anleu, R., McNab. R., and D.H. Thornton**. 2021. When waterholes get busy, rare interactions thrive: Photographic evidence of a jaguar (Panthera onca) killing an ocelot (Leopardus pardalis). Biotropica 53:367-371

Olson, L., Bjornlie, N., Hanvey, G., Holbrook, J.D., Ivan, J.S., Jackson, S., Kertson, B., King, T., Lucid, M., Murray, D., Naney, R., Rohrer, J., Scully, A., Thornton, D., Walker, J. and J.R. Squires. 2021. Regional transferability and data efficiency within Species Distribution Modelling: an evaluation for a specialist carnivore, the Canada Lynx. Ecology and Evolution 11:1167-1690.

Turcios-Casco, M.A., Avila-Palma, H.D., Martinez, M., Trejo-Ordonez, E.J., Meza-Flores, D.E., Castaneda, F., King1, T., Thornton, D.H., and M. Superina. 2021. The xenarthrans of Honduras: new records, main threat, and comments on their conservation status. Edentata. 21:13-28.

Martinez, W.E., Reyna-Hurtado, R.A., Naranjo, E.J., Thornton, D.H., Cal, R.N., and Figueroa, O.A. 2021. Occupancy rate and observations of Baird’s tapir (Tapirella bairdii) near waterholes in the Maya forest corridor, Belize. Therya 12:37-43.



Thornton, D.H., Reyna, R., Perera-Romero1, L., Radachowsky, J., Hidalgo-Mihart, M.G., Garcia, R., McNab, R., Mcloughlin, L., Foster, R., Harmsen, B., Moreira-Ramírez, J.F., Diaz-Santos, F., Jordan, C., Salom- Pérez, R., Meyer, N., Castañeda, Valle, F.A.E., Santizo, G.P., Amit, R., Arroyo-Arce, S., Thomson, I., Moreno, R., Schank, C., Arroyo-Gerala, P., Bárcenas, H.V., Brenes-Mora, E., Calderón, A.P., Cove, M.V., Gomez-Hoyos, D., González-Maya, J., Guy, D, Jiménez, G.H., Hofman, M., Kays, R., King1, T., Menjivar, M.A.M., de la Maza, J., León-Pérez, R., Ramos, V.H., Rivero, M., Romo-Asunción, S., Juárez-López, R., Cruz, A-J., de la Torre, J.A., Towns, V., Schipper, J., Reyes, H.O.P., Artavia, A., Hernández-Perez, E., Martínez, W.,Urquhart, G.R., Quigley, H., Pardo, L.E., Sáenz, J.C., Sanchez, K., and J. Polisar. 2020. Precipitous decline of white-lipped peccary populations in Mesoamerica. Biological Conservation 242:108410. [IF=5.0/NA/NA]

Thornton, D.H., Branch, L.C., and D.L. Murray. 2020. Distribution and connectivity of protected areas in the Americas facilitates transboundary conservation. Ecological Applications 30:e02027 

Thornton, D.H. and M.J.L. Peers. 2020. Chapter 8. Species Distribution Modeling. Population Ecology in Practice (Murray, D.L. & Chapron, G., eds). Wiley-Blackwell, London.

Murray, D.L., Bastille-Rousseau, G., Hornseth, M., Row, J. F , and D.H. Thornton. 2020. Chapter 3. From research hypothesis to model selection: A strategy toward robust inference in population ecology. Population Ecology in Practice (Murray, D.L. & Chapron, G., eds). Wiley-Blackwell, London.

Olsoy1, P.J., Forbey, J.S., Shipley, L.A., Rachlow, J.L., Robb, B.C., Nobler, J.D., and D.H. Thornton**. 2020.  Mapping foodscapes and sagebrush morphotypes with unmanned aerial systems for multiple herbivores. Landscape Ecology 35:921-936.

King1, T., Vynne, C., Miller, D., Fisher, S., Fitkin, S., Rohrer, J., Ransom, J., and D.H. Thornton**. 2020. Will Lynx Lose Their Edge? Canada Lynx Occupancy in Washington. Journal of Wildlife Management 84:705-725.



Thornton, D.H., and L.C. Branch. 2019. Transboundary mammals in the Americas: asymmetries in protection challenge climate change resilience. Diversity and Distributions 25:674-683.

Thornton, D.H., King1, T., Scully2, A., and D.L. Murray. 2019. Reassessing the success of experts and non-experts in correctly differentiating between closely related species from camera trap images: A reply to Gooliaff and Hodges. Ecology and Evolution 9:6172-6175.

Ebenhoch1, K., D.H. Thornton**, Shipley, L., Manning, J., White, K. 2019. Effects of post-release movements on survival of translocated sage-grouse. Journal of Wildlife Management 83: 1314-1325.

Santos, P. M., A. Bocchiglieri, A. G. Chiarello, A. P. Paglia, A. Moreira, A. C. de Souza, A. M. Abba, A. Paviolo, A. Gatica, A. Z. Medeiro,….D.H. Thornton…et al. 2019. NEOTROPICAL XENARTHRANS: a data set of occurrence of xenarthran species in the Neotropics. Ecology 100:e02663.



Thornton, D.H., Wirsing, A.W., Lopez-Gonzalez, C.L., Squires, J.R., Fisher, S., Larsen, K.W., Peatt, A., Scrafford, M.A., Moen, R.A., Scully, A.E., King, T.W., and D.L. Murray. 2018. Asymmetric cross-border protection of peripheral transboundary species. Conservation Letters 11:e12430.

Thornton, D.H., Scully1, A., King1, T., Fisher, S., Fitkin, S., and J. Rohrer. 2018. Hunting associations of badgers and coyotes revealed by camera trapping. Canadian Journal of Zoology 96:769-773.

Olsoy1, P.J., Shipley, L.A., Rachlow, J.L., Forbey, J.S., Glenn, N.F., Burgess, M.A., and D.H. Thornton**. 2018. Unmanned aerial systems measure structural habitat features for wildlife across multiple scales. Methods in Ecology and Evolution. 9:594-604.

Scully1, A.E., Fisher, S., Miller, D.A.W., and D.H. Thornton**. (This article was Editor’s Choice for the issue) 2018. Influence of biotic interactions on Canada lynx (Lynx canadensis) distribution at their southern range edge. Journal of Mammalogy 99:760-772.

Milling, C.R., J.L. Rachlow, P.J. Olsoy, M.A. Chappell, T.R. Johnson, J.S. Forbey, L.A. Shipley, D.H. Thornton. 2018. Habitat structure modifies microclimate: an approach for mapping fine-scale thermal refugia. Methods in Ecology and Evolution 9:1648-1657.



King2, T.W., Salom-Perez, R., Shipley, L.A., Quigley, H.B., and D.H. Thornton**. 2017. Ocelot latrines: communication centers for Neotropical Mammalia. Journal of Mammalogy. 98: 106-113. 

Otis, J-A, Thornton, D.H., Rutledge, L, and D.L. Murray. 2017. Ecological niche differentiation across a wolf-coyote hybrid zone in eastern North America. Diversity and Distributions. 23:529-539.

Camp, M., Shipley, L., Johnson, T., Olsoy, P., Crowell, M., Forbey, J., Rachlow, J., and D.H. Thornton. 2017. The balancing act of foraging: mammalian herbivores trade-off multiple risks when selecting food patches. Oecologia 185: 537-549.

Gelin, M.L., Branch, L.C., Thornton, D.H., Novaro, A.J., Gould, M.J., and Caragiulo, A. 2017. Response of pumas (Puma concolor) to migration of their primary prey in Patagonia.  PlosOne 12: e0188877. [IF=3.3/2/2]

Feldman, R. Peers, M.J., Pickles, R.S.A., Thornton, D.H., and D.L. Murray. 2017. Climate driven niche divergence among host species affects range-wide patterns of parasitism. Global Ecology and Conservation 9:1-10.

Murray, D.L., Peers, M.J.L., Majchrzak, Y.N., Wehtje, M., Ferreira, C., Pickles, R.S.A., Row, J.R., and D.H. Thornton.  2017. Continental divide: Predicting climate-mediated fragmentation and biodiversity loss in the boreal forest. PlosOne 12:e0176706.

Hidalgo-Mihart, M.G., Jesús de la Cruz, A., Juárez-Lopez, R., Contreras-Moreno, F.M., Bravata de la Cruz, C., Hernández-Lara, C., Pérez-Solano, L.A., Kohler González, J.M., Friedeberg, D., and D.H. Thornton. 2017. Medium and large –sized mammal inventory in the wetlands of Laguna de Terminos and Pantanos de Centla, México. Checklist: Journal of Biodiversity Data 13:711-726.

Juárez-López, R., Pérez-López, M., Bravata-de la Cruz, Y., Jesus-de la Cruz, J., Contreras-Moreno, F.M., Thornton, D.H., Hidalgo-Mihart, M.G. 2017. Range extension of the northern naked-tailed armadillo (Cabassous centralis) in southern Mexico.  Western North American Naturalist 77:398-403.



Thornton, D.H., Zeller, K., Rondinini, C., Boitani, L., Crooks, K., Burdett, C., Rabinowitz, A., and H. Quigley. 2016. Assessing the umbrella value of a range-wide conservation network for jaguars (Panthera onca). Ecological Applications 26:1112-1124.

Olsoy1, P.J., Zeller, K.A., Hicke, J.A., Quigley, H.B., Rabinowitz, A.R., and D.H. Thornton**. 2016. Quantifying the effects of deforestation and fragmentation on a range-wide conservation plan for jaguars. Biological Conservation 203:8-16.

Peers1, M.J., Thornton, D.H., Majchrzak, Y.N., Bastille-Rousseau, G., and Murray, D.L. 2016. De-extinction potential under climate change: extensive mismatch between historic and future habitat suitability for three candidate birds. Biological Conservation 197:164-170.



Guillaumet, A., Bowman, J., Thornton, D.H., and D.L. Murray. 2015. The influence of coyote on Canada lynx populations assessed at two different spatial scales. Community Ecology 16:135-146.

Thornton, D.H., and Pekins, C. 2015. Spatially-explicit capture recapture analysis of bobcat density in central Texas: implications for mesocarnivore monitoring. Wildlife Research 42:394-404.

Murray, D.L., Majchrzak, Y.N., Peers, M.J.L., Wehtje, M., Ferreira, C., Pickles, R.S.A., Row, J.R., and D.H. Thornton. 2015. Potential pitfalls of private initiatives in conservation planning: A case study from Canada’s boreal forest. Biological Conservation 192: 174-180.

Wang, Y., Thornton, D.H., Wang, S., Ding, P. 2015. Ecological correlates of vulnerability to fragmentation in forest birds on inundated subtropical land-bridge islands. Biological Conservation 191:251-257.



Thornton, D.H., and D.L. Murray. 2014. Influence of hybridization on niche shifts in expanding coyote populations. Diversity and Distributions 20: 1355-1364.

Row, J.R., Wilson, P.J., Gomez, C, Koen, E.L., Bowman, J., Thornton, D., and D.L. Murray. 2014. The subtle role of climate change on population genetic structure in Canada lynx. Global Change Biology 20: 2076-2086.

Thornton, D.H., and R. Fletcher. 2014. Body size and spatial scales in avian response to landscape structure: a meta-analysis. Ecography 37:454-463.

Peers2, M.J.L., Wehtje, M., Thornton, D.H. and D.L. Murray. 2014. Prey switching as a means of enhancing persistence in predators at the southern range edge. Global Change Biology 20:1126-1135.



Peers2, M.J.L., Thornton, D.H. and D.L. Murray. 2013. Evidence for large-scale effects of competition: niche displacement in carnivores. Proceedings of the Royal Society B 280: 20132495.

Pickles, R.S.A., Thornton, D.H., Marques1, A., D. L. Murray. 2013. Predicting shifts in parasite distribution with climate change: a multitrophic level approach. Global Change Biology 19: 2645-2654.

Thornton, D.H., A.J. Wirsing, J.D. Roth, and D.L. Murray 2013. Habitat quality and population density drive occupancy dynamics of snowshoe hare in variegated landscapes. Ecography 36: 610-621.



Thornton, D.H. A.J. Wirsing, J.D. Roth, and D.L. Murray. 2012. Complex effects of harvest and site preparation on snowshoe hare abundance on a patchy forest landscape. Forest Ecology and Management 280: 132-139.

Thornton, D.H., L.C. Branch, and M.E. Sunquist. 2012. Response of large galliforms and tinamous (Cracidae, Phasianidae, Tinamidae) to habitat loss and fragmentation in northern Guatemala. Oryx 46: 567-576.

Peers1, M.J.L., Thornton, D.H., and D.L. Murray. 2012. Reconsidering the specialist-generalist paradigm in niche breadth dynamics: Canada lynx and bobcats. PLoSOne. 7(12): e51488. doi:10.1371/journal.pone.0051488



Thornton, D.H., L.C. Branch, and M.E. Sunquist. 2011. The relative influence of habitat loss and fragmentation: Do tropical mammals meet the temperate paradigm? Ecological Applications 21(6):2324-2333.

Thornton, D.H., L.C. Branch, and M.E. Sunquist. 2011. Passive sampling effects and landscape location alter associations between species traits and response to fragmentation. Ecological Applications: 21(3):817-829.

Thornton, D.H., L.C. Branch, and M.E. Sunquist. 2011. The influence of landscape, patch, and within-patch factors on species presence and abundance: a review of focal patch studies. Landscape Ecology: 26:7-18.



Thornton, D.H., M.E. Sunquist, and M.B. Main. 2004. Ecological separation within newly sympatric populations of coyotes and bobcats in southern Florida. Journal of Mammalogy 85(5):973-982.